良質な牛乳生産を目指して、反芻動物の乳房炎を防除するために乳房の自然免疫に着目している。乳房内の抗菌物質（defensin, S100, cathelicidin, lactoferrin）等の免疫物質を最大限利用して、感染を防御する手法を検討している。

My study focuses on innate immune function in mammary glands to prevent and treat mastitis. I try to
establish the appropriate methods to maximize the amount of antimicrobial components（defensin, S100, cathelicidin, lactoferrin）in milk. Mastitis caused by the bacterial components translocated from uterus was identified.

keywords：mastitis, mammary gland, antimicrobial component, cow, goat, milk

1. Colchicine can keep the viability of bacteria in mastitic milk by preventing leukocyte phagocytosis in dairy cow and goat. Front Vet Sci 11:1469586.（2024）
2. Effect of low milking frequency on the concentration of antimicrobial proteins in goat milk. Vet Immunol Immunopathol 272: 110774（2024）
3. Interleukin 1 receptor antagonist（IL-1ra）production and expression dynamics in mammary glands after lipopolysaccharide infusion. Anim Sci J 96: e70047（2025）

世界に広がる塩害土壌でも栽培可能な作物品種の育成や遺伝子組換えによる耐塩性作物の創出を行っています。また、植物の生育を促進する善玉菌の単離とその実用化技術、肥料資源の節約技術の開発を行っています。

Our research focuses on molecular physiological analysis of environmental stress（high salinity, nutrient deficiency, etc.）tolerance in higher plants for sustainable crop production. We also study plant growth promoting bacteria to increase crop production through improving bacterial fitness in the rhizosphere and identifying bioactive compounds useful for crop growth.

keywords：sustainable crop production, rice, salinity stress, nutrient deficiency, plant nutrition, plant growth promoting bacteria

1. A chromosome-level genome sequence reveals regulation of salt stress response in Mesembryanthemum crystallinum. Physiol Plant 177: e70057（2025）
2. Exogenous riboflavin（vitamin B2）application enhances salinity tolerance through the activation of its biosynthesis in rice seedlings under salinity stress. Plant Sci 339: 111929（2024）
3. Potassium transporter OsHAK17 may contribute to saline-alkaline tolerant mechanisms in rice（Oryza sativa）. J Plant Res 137: 505-520 （2024）

瀬戸内海の重要種を対象に、種苗放流や資源管理によって、資源を増殖し、持続的に利用するための研究を行っている。特に、クロダイは研究の代表種で、最近では産卵生態や移動回遊の解明に取り組んでいる。

My current research focuses on sustainable use of commercially important fish by using stock enhancement and resource management. Black sea bream is representative fish of my research, and I want to pursue studies to clarify all of the mysteries of this fish as my life work.

keywords：fish, black sea bream, stock enhancement, resource management, sustainable use

1. The role of the isolation of the marginal seas during the Pleistocene in the genetic structure of black sea bream Acanthopagrus schlegelii （Bleeker, 1854）in the coastal waters of Japan. PeerJ 9（24）:e11001（2021）
2. Estimating the spawning season of black sea bream Acanthopagrus schlegelii in Hiroshima Bay, Japan, based on temporal variation in egg density. Fish Sci, 86:645-653（2020）
3. Infestation of the parasitic isopod Mothocya parvostis on Juveniles of the black sea bream Acanthopagrus schlegelii as an optional intermediate host in Hiroshima Bay. Zool Sci,1-10（2020）

飼料資源の有効利用（副産物飼料）、環境負荷の低減（ウシからの温室効果ガス）、栄養代謝調節（アミノ酸代謝）などを目標にして、反芻家畜（ウシ、ヤギ、ヒツジ）の飼料と栄養代謝に関する研究を行っている。

In order to establish sustainable and healthy ruminant production systems, I am studying by-product feed utilization, mitigation of enteric methane emission from ruminants, and regulation of amino acid metabolism in cattle and sheep by the natural functional compounds in forage.

keywords：forage, digestion, metabolism, protein, amino acid, cow, sheep, milk, meat.

1. Phytol supplementation alters plasma concentrations of formate, amino acids, and lipid metabolites in sheep. Animal 15:100174.（2021）
2. Effects of ensiling treatment for tuber crop forages and grain source on carbohydrate digestion, nitrogen utilization, and urea metabolism in sheep. Anim Feed Sci Technol 243: 140-149.（2018）
3. Effects of nitrogen fertilizer and harvesting stage on photosynthetic pigments and phytol contents of Italian ryegrass silage. Anim Sci J 88: 1513 – 1522.（2017）

単細胞藻類を専門とし、その環境適応性、光合成機能等を培養実験・フィールド調査両面から探っています。彼らが支える生態系（海洋の生物生産システム、マングローブ・サンゴ礁生態系）の保全に役立てたいと考えています。

Microalgae, or single-cell algae, especially their diversity, environmental adaptation and photosynthetic
mechanisms are investigated both from field studies and laboratory culture experiments. Goal of the study is to apply these knowledge for conserving characteristic ecosystems（e.g. coastal biological production, mangrove ecosystem, coral reef ecosystem, etc）whose basis are microalgae.

keywords：microalgae, aquatic ecosystem, coast, mangrove, coral reef

1. Fecal pellets of giant clams as a route for transporting Symbiodiniaceae to corals/ PLoS ONE/2020
2. Zooxanthellal genetic varieties in giant clams are partially determined by species-intrinsic and growth-related characteristics/ PLoS ONE/ 2017
3. Aplication of a pulse-amplitude-modulation（PAM）fluorometer reveals its usefulness and robustness in the prediction of Karenia mikimotoi blooms: A case study in Sasebo Bay, Nagasaki, Japan/ Harmful Algae/ 2017

サンゴ礁などリーフ環境に棲む魚類の採餌生態や産卵生態に注目するフィールド調査から、生存戦略や繁殖戦略を解明する研究を進めている。

Behavioral ecology of reef fishes focusing on survival and reproductive strategies（e.g. foraging and mating tactics, social and mating system, and sex change patterns）by using the method of animal sociology including field observation and demographic survey in nature.

keywords：underwater observation, field survey, function of behaviors, intra-and interspecific relationships, individual-level data, reproductive success

1. Reversed sex change in the haremic protogynous hawkfish Cirrhitichthys falco in natural conditions. Ethology 118: 1-9（2012）
2. Effect of changing harem on timing of sex change in female cleaner fish Labroides dimidiatus. Anim Behav 62: 251-257（2001）
3. Alternative spawning tactics of female angelfish according to two different contexts of sex change. Behav Ecol 8: 372-377（1997）

乳牛の哺育期および分娩移行期など栄養管理に起因する疾病リスクの高いステージに着目し、被害軽減に向けた栄養生理学的研究に取り組んでいる。初乳給与と消化管ホルモン、小腸発達との関連性などを明らかにしている。

My work focuses on the nutrition and metabolism at transition periods for weaning and calving in dairy
cattle, especially the relationship between dietary nutrition and metabolic hormone action.

keywords：gastro-intestinal hormone, dairy cow, dairy calf, nutritional management

1. Effects of medium-chain fatty acids and tributyrin supplementation in milk replacers on growth performance, blood metabolites, and hormone concentrations in Holstein dairy calves. J. Dairy Sci. 106, 4599–4607.（2023）
2. Effect of kraft pulp inclusion in calf starter on performance, health, and plasma concentration of glucagon-like peptide 2 in calves. J. Dairy Sci. 106, 4443–4453.（2023）
3. Effects of rumen-protected glutamate supplementation during the periparturient period on digestibility, inflammation, metabolic responses, and performance in dairy cows. J. Dairy Sci. 105, 3129–3141.（2022）

植物の表皮細胞分化に関わる転写因子の研究を行っている。表皮細胞は根毛のような特殊な器官に分化する。分化制御ネットワークの解明により、植物の形態形成や細胞分化の仕組みを分子レベルで理解する事を目指す。

The aim of our study is to elucidate the mechanism of epidermal cell differentiation in plants, by using epidermal cells from specialized organs, such as root hairs. By focusing on the function of transcription factors, we aim to understand plant morphogenesis and cell differentiation at the molecular level.

keywords：Arabidopsis, root hair, transcription factor

1. Effect of shading on trichome formation and CAPRICE-like gene expression in tea（Camellia sinensis var. sinensis）leaves. Sci. Hortic. 330: 113049（2024）
2. Effect of phosphorus deficiency on the expression and protein localization of ENHANCER OF TRY AND CPC3 gene in Arabidopsis roots. J. Plant Biochem. Biotech. 33: 260-264（2024）
3. Complementation and protein localization analyses of R3 MYBs in an Arabidopsis caprice mutant. Plant Biotech. 40: 99-103（2023）

瀬戸内海を中心に、日本沿岸における魚介類の資源生態を研究している。特に稚魚期の摂食や成長に関する研究のほか、地球温暖化など環境の変化に対する水産資源の応答に関する研究を進めている。

I am studying fisheries biology in Japan, especially in the Seto Inland Sea, to contribute to sustainable
coastal fisheries. My research interests are mainly in the life history traits of commercially important species, predator-prey interactions, and the response of fishes to climate change.

keywords：aquatic ecology, fisheries, life history, thermal biology, tidal flat

1. Effects of hook size on the catch variation, size selectivity, and bycatch of the bottom set longline fisheries targeting nemipterid fishes in the Philippines. Sci Rep 15: 1379（2025）
2. Effects of constant and variable salinity regimes on the occurrence, feeding, and growth of two juvenile flatfishes（Pseudopleuronectes yokohamae and Platichthys bicoloratus）. Estuar Coast Shelf Sci 305: 108872（2024）
3. Potential impact of predation by larval Spanish mackerel on larval anchovy in the central Seto Inland Sea, Japan. Deep-Sea Res. II 208: 105272（2023）

ゲノム情報をもとに日本鶏をはじめとした家畜・家禽の起源や世界伝播プロセス、そして多様な表現型を創出する遺伝的基盤を解明し、さらなる育種を進めていくことを目指しています。
Based on genome information, we aim to elucidate the domestication origins, the global propagation processes, and the genetic basis for the diverse phenotypes of Japanese native chickens and other livestock, in order to promote further breeding.

keywords：genome, evolution, breeding

1. Phylogenomics and morphology of extinct paleognaths reveal the origin and evolution of the ratites. Current Biology, 27:68-77（2017）
2. Paleogenomics reveals independent and hybrid origins of two morphologically distinct wolf lineages endemic to Japan. Current Biology 32:2494-2504（2022）
3. Complete mitochondrial genome analysis clarifies the enigmatic origin of Haplogroup D in Japanese native chickens. The Journal of Poultry Science, 59: 316-322（2022）

海洋の生物と環境の相互作用を研究している。生物から環境への作用は、生物による海洋への炭素隔離過程を研究している。環境から生物への作用は、二酸化炭素の増加に対する海洋生態系の応答を主な研究課題である。

We study how marine organisms interact with the environment, aiming to understand two main effects:
how organisms influence the environment, like carbon sequestration, and how the environment affects organisms, such as ecosystem responses to rising CO₂ levels.

keywords：carbon sequestration, rising CO₂ level

1. Massive loss and microbial decomposition in reproductive biomass of Zostera marina. Est Coast Shelf Sci 275:107986（2022）
2. Ocean acidification increases phytobenthic carbon fixation and export in a warm-temperate system. Est Coast Shelf Sci 250:107113（2021）
3. Contribution of macroalgae to coastal dissolved organic matter pool. Est Coast Shelf Sci 129:77-85（2013）

農業生産において循環型イニシアチブを確立し、応用可能な農産物を生産します。特に米の副産物を活用して食品、医薬品、肥料を開発します。また、突然変異および交配法を用いて高収量の米品種を作出します。

We are establishing a circular initiative in agricultural production to generate highly applicable agricultural products. Our focus is on the utilization of agricultural by-products, particularly from rice production, for developing foods, pharmaceuticals, and fertilizers. Additionally, we are applying mutation and crossbreeding methods to create high-yield rice varieties.

keywords：circular agricultural production, biochemistry, natural products, biological activities, plant breeding, genetic engineering

1. Cytotoxic mechanism of momilactones A and B against acute promyelocytic leukemia and multiple myeloma cell lines. Cancers 14:4848（2022）
2. Uniparental Inheritance of Salinity Tolerance and Beneficial Phytochemicals in Rice. Agronomy-Basel 10:1032（2020）
3. Decomposition of allelopathic plants in soil. J Agron Crop Sci 191:162（2005）

環境分析化学を活用して、幅広い水環境問題の解決に挑戦しています。特にリサイクル材料を有効活用した沿岸環境の修復、生活・産業排水処理技術の開発、水環境を評価するための分析法の開発について研究しています。

I am an aquatic environmental scientist with a broad perspective of wastewater treatment, restoration of coastal seas etc. based on my analytical chemistry background. My specialties and skills include（1）Coastal environment restoration using industrial recycled materials, （2）Development of analytical methods to evaluate aquatic environment.

keywords：aquatic environment, environmental analytical chemistry, environmental remediation, SDGs, wastewater treatment

1. Identification of sulfur species in coastal marine sediments collected from the Seto Inland Sea, Japan in summer, Estuar. Coast. Shelf Sci. 313:109130（2025）
2. Adsorptive removal of phosphate from aqueous solutions using iron–lanthanum-doped foam glass adsorbent, Water Environ. Res. 97: e70025（2025）
3. Enhanced growth of benthic microalgae by tablet from liquid dairy cattle manure-based anaerobic digestate, Chemosphere 363:142943 （2024）

子どもができる仕組みについて、代謝と内分泌に焦点を当て、モデル動物を用いた基礎研究から、家畜を用いた実用化研究まで行っている。この成果を基盤にした“効率よく家畜生産ができる繁殖技術開発”を目指している。

Our aim is to develop the reproductive technology in animal production. To achieve this aim, we are focusing on mammalian reproductive biology of both female and male, such as ovary, oocyte, testis and sperm, especially focusing on metabolism, endocrine and immune system.

keywords：animal reproduction, reproductive technology, endocrine, metabolism

1. A simple sperm-sexing method that activates TLR7/8 on X sperm for the efficient production of sexed mouse or cattle embryos. Nature protocols,（2020）
2. Activation of Toll-like receptor 7/8 encoded by the X chromosome alters sperm motility and provides a novel simple technology for sexing sperm. PLoS Biol.,（2019）
3. The acceleration of reproductive aging in Nrg1flox/flox; Cyp19‐Cre female mice. Aging cell, 16, 1288-1299.（2017）

沿岸生態系の生物多様性の維持に欠かせない藻場や海藻養殖で顕在化する、気候変動の影響に注目している。海藻類の系統分類や海藻相研究を行う一方、食用海藻や石灰藻について実験下での生育特性を把握してきた。

Seaweeds benefit human life by providing food and indirectly by maintaining biodiversity in coastal ecosystems. Focusing on the impacts of climate change on seaweed aquaculture and seaweed forests, I have examined molecular phylogeny and ecophysiology of key species of macroalgae and conducted empirical research relevant for aquaculture.

keywords：coralline algae, edible seaweeds, optimum growth conditions, rhodolith, taxonomy

1. Coralline red algal species diversity at a shallow rhodolith bed in warm-temperate Japan, including two new species of Roseolithon（Hapalidiales, Corallinophycidae）. Phycologia 63: 520–533（2024）
2. Lithophyllum nagaokaense sp. nov.（Corallinales, Corallinophycidae, Rhodophyta）: A new rhodolith‐forming non‐geniculate coralline alga from Japan. Phycological Research 72: 167-179（2024）
3. Effects of seawater temperature and irradiance on the growth of sporelings of Amphiroa cf. zonata and Corallina berteroi（Corallinales, Rhodophyta）in Japan. Phycologia 62: 585-592（2023）

主にニワトリをモデル動物として用い、本能行動（摂食行動・飲水行動・攻撃行動など）の脳内メカニズムについて、間脳視床下部を中心に研究しています。

Research on the brain mechanisms of innate behavior such as feeding, drinking, and aggressive behavior in chickens.

keywords：feeding behavior, drinking behavior, aggressive behavior, hypothalamus, chicken

1. Screening of the behavioral tests for monitoring agonistic behavior of layer chicks. J.Poult.Sci. 54:296-302.（2017）
2. Blood testosterone concentration and testosterone-induced aggressive behavior in male layer chicks: Comparison between isolated- and grouped-raising. J.Poult.Sci. 56:290-297.（2019）
3. Effects of testicular and non-testicular testosterone on territorial and isolation-induced aggressive behavior of male layer chicks. J.Poult. Sci. 57:236-240.（2020）

社会ネットワーク解析を通じた乳牛（搾乳牛、育成牛）の社会行動、個体間の親和的、敵対的関係の総合的解析を行い、家畜福祉と高い生産性の両方を実現する飼養管理システムの構築を目指す。

The aim of my study is construction of management system achieving higher animal welfare and productivity by the analysis of social behavior, affinity and agonistic relationships between dairy cows (lactating cows and raising cows) through social network analysis.

keywords：social network analysis, dairy cows, milk production, social behavior

1. Effect of relationships among clinical mastitis incidence, reproductive performance, and culling rate on the lifetime of dairy cows at Hiroshima University Farm./Anim. Sci. J. /2021
2. Changes of photosynthetic pigments and phytol content at different levels of nitrogen fertilizer in Italian ryegrass fresh herbage and hay./ Grassl. Sci./2022
3. A comparison of plasma glucose and oxidative status in lactating dairy cows in summer and autumn/Anim. Sci. J./2016

釣り餌として利用される多毛類やエビ類を対象とした生態学的研究、とくに外来種としての分布状況や生物多様性に及ぼす影響、および希少在来種の保全と有効活用を目的とした生活史の解明について研究している。

My research interests are ecological researches on aquatic organisms, especially understanding of life history aimed at the dispersal mechanism of alien species and the conservation of threatened species.

keywords：benthos, fishing bait, freshwater, intertidal flat, polychaete, shrimp

1. Appearance of exotic shrimp Palaemon sinensis (Sollaud, 1911) and other freshwater shrimps before and after the 2018 extreme flood in western Japan. BioInvasions Rec. 13: 183-194 (2024)
2. Host selection and potential predation in host-parasite interaction between the isopod Tachaea chinensis and freshwater species. Int J Parasitol: Parasites Wildl. 21: 59-68 (2023)
3. Population genetics of the non-native frechwater shrimp Palaemon sinensis (Sollaud, 1911) in Japan based on mitochondrial 16S rRNA sequence analysis. Aquat. Invasions. 16: 710-720 (2021)

反芻動物乳房への細菌感染成立機序解明を基盤として、乳房炎制御による食（乳）の安定供給と生乳衛生の向上による食（乳）の安全に貢献することを目指している。また、乳房炎迅速診断技術開発も行っている。

The aim of my studies is to contribute food（milk）supply and food（milk）safety by elucidating mechanisms of bacterial intramammary infection in dairy animals.

keywords：dairy animals, intramammary infection, zoonosis, diagnosis

1. Seasonal variations in the concentration of antimicrobial components in milk of dairy cows. Anim Sci J. 91（1）.（2020）
2. Outcome prediction from the first examination in clinical mastitis using ultrasonography in dairy cows. Anim Sci J. 91（1）.（2020）
3. Rapid determination of pathogens in mastitic milk of dairy cows using Gram staining. J Vet Med Sci. in press.（2022）

鳥類とほ乳類の生殖幹細胞を対象として、培養・凍結保存・移植等の操作技術を開発・改善し、動物の遺伝資源を細胞レベルで保存すること、ヒト不妊治療技術を開発することに挑戦している。

My research group aims to develop culture, cryopreservation and transplantation of germ cells including primordial germ cells and spermatogonial stem cells in birds and mammals, and to apply germ cell manipulation technologies from conservation of biological diversity to human infertility treatment.

keywords：germ cells, chicken, mouse, genetic conservation, infertility treatment

1. Transient suppression of transplanted spermatogonial stem cell differentiation restores fertility in mice. Cell Stem Cell 28:1-14.（2021）
2. Efficient system for preservation and regeneration of genetic resources in chicken: concurrent storage of primordial germ cells and live animals from early embryos of a rare indigenous fowl（Gifujidori）. Reprod Fertil Dev 22:1237-1246.（2010）
3. Germline replacement by transfer of primordial germ cells into partially sterilized embryos in the chicken. Biol Reprod 83:130-137.（2010）

持続可能な植物生産のために、有用資源の有効活用と土壌中での養分の動態および植物による利用性の改善に関する研究を行っています。堆肥などの有機物施用による作物（イネなど）の生産性や養分利用効率を解析しています。

Aiming for sustainable plant production, we are conducting researches on the effective utilization of useful resources, dynamics of nutrients in soils, and improvement of their utilization by plants. We analyze productivity and nutrient use efficiency of crops（such as rice）by applying organic matter such as compost.

keywords：plant production, soil, organic matter, nutrient cycling

1. Shoot and root responses to low phosphorus and their genotypic variability in selected cultivars of Japanese core collections of maize and soybean. Soil Sci. Plant Nutr. 70: 100-113 (2024)
2. Phytate degradation by fungi and bacteria inhabiting sawdust and coffee residue composts. Microbes Environ. 28: 71-80 (2013)
3. Isolation and characterization of cellulose-decomposing bacteria inhabiting sawdust and coffee residue composts. Microbes Environ. 27: 226-233 (2012) 

ニワトリの健康と安全な卵の生産を目指して、粘膜バリア機能の強化と産卵機能改善に関する研究に取組んでいる。特に、腸内環境を中心として、肝臓や脳、卵管といった周辺の組織とのネットワークに注目している。

I work on the research about enhancement of mucosal barrier function and improvement of egg production for healthy and secure poultry production of chickens. In particular, I focus on the intestinal environment and the network between intestine and surrounding organs such as liver, brain and reproductive organs.

keywords：avian, mucosal immunity, digestive tract cellular immunity, reproductive organ, egg laying function, probiotics

1. Slight Disruption in Intestinal Environment by Dextran Sodium Sulfate Reduces Egg Yolk Size Through Disfunction of Ovarian Follicle Growth. Front Physiol. 11:607369.（2021）
2. Intestinal inflammation induced by dextran sodium sulphate causes liver inflammation and lipid metabolism disfunction in laying hens. Poult Sci. 99:1663-1677.（2020）
3. Effects of avian infectious bronchitis virus antigen on eggshell formation and immunoreaction in hen oviduct. Theriogenology. 81: 1129-1138.（2014）

持続的な漁業生産可能な海洋環境を維持・管理するために、コンピュータによるデータ解析、生態系モデル解析や海洋調査により沿岸海域の物質循環機構と低次生物生産構造を明らかにするための研究を行っている。
The main purpose of my study is to maintain and manage the marine environment for sustainable fishery production. For the purpose, I perform studies about the material circulation system in the coastal seas by computer data analysis, numerical ecosystem model and field observations.

keywords：sustainable fishery production, computer data analysis, numerical ecosystem model

1. Primary Production and Physical Structure of the Seto Inland Sea, Japan / Bull. Coastal Oceanogr.（1997）
2. Analysis of Primary Production in the Seto Inland Sea, Japan, Using a Simple Ecosystem Model/ J. Oceanogr. 54: 123-132.（1998）
3. Optical Properties of Sea Water in Hiroshima Bay, Japan / Oceanography in Japan（1997）

生物学的心理学を専門とする。動物の心を生物学的な機能のひとつとして考える。脳を含む身体という有形物から、心という形のないものがいかにして生じるのかを探究している。

Biological psychology and behavioral neuroscience. Studying animal mind as a biological function. My research involves quantitative observation of behavior and manipulation of underlying neural activities to evaluate emotional states of animals.

keywords：behavior, biological psychology, brain, emotion

1. Recording the ventilation activity of free-swimming zebrafish and its application to novel tank tests. Physiology & Behavior, 244, 113665 （2022）
2. Measurement of emotional states of zebrafish through integrated analysis of motion and respiration using bioelectric signals. Scientific Reports 11:187（2021）
3. Relationship between brain morphology and life history in four bottom-dwelling gobiids. Zoological Science, 37, 168-176（2020）

海産無脊椎動物の初期生活史に関して分類・発生・生態の観点から研究している。最近は養殖候補種である甲殻類や棘皮動物の安定的な種苗生産や畜養の技術開発に取り組んでいる。
My main research interest is the early life history of marine invertebrates particularly from the viewpoints of taxonomy, developmental biology, and ecology. Applied aquaculture sciences of crustaceans and echinoderms are also the research topics that we focus on.

keywords：lobster, larvae, jellyfish, hatchery, land-based aquaculture

1. The final phyllosoma, nisto, and first juvenile stages of the slipper lobster Petrarctus brevicornis（Holthuis, 1946）（Decapoda: Achelata: Scyllaridae）. J Crustac Biol 40: 237–246.（2020）
2. Culture of slipper lobster larvae（Decapoda; Achelata; Scyllaridae）fed jellyfish as food. In Lobsters: Biology, Fisheries and Aquaculture. Springer Singapore, Singapore, pp. 519–540.（2019）
3. Amended larval recruitment model for the Japanese spiny lobster Panulirus japonicus from new larval records and population genetic data in Taiwan. J Oceanogr 75: 273–282.（2019）

反芻家畜（乳牛・肉牛）の健全性を担保しつつ生産性を向上させることを目標に、生理・内分泌機構や腸内環境に着目した研究を行っている。

Research focusing on physiological and endocrine mechanisms and intestinal environment to improve health and productivity of ruminants (dairy and beef cattle).

keywords：ruminants, endocrinology, physiology, hormones, gut-derived peptides, nutrition, intestinal environment

1. Effect of extended colostrum feeding on plasma glucagon-like peptide-1 concentration in newborn calves. J Dairy Sci: 4619-4627 (2019)
2. Effect of kraft pulp inclusion in calf starter on performance, health, and plasma concentration of glucagon-like peptide 2 in calves. J Dairy Sci 4443-4453 (2023)
3. Development of a novel feeding method for Japanese black calves with thermophile probiotics at postweaning. J Appl Microbiol 3870-3882 (2022)

ゲノムデータを活用して複雑な表現型の背後にある遺伝的メカニズムの解明を目指している。またウイルスゲノム解析を通した疫学的研究や家畜のゲノム育種研究なども行っている。

I aim to uncover the genetic mechanisms of complex traits using genome big data, grounded in evolutionary biology and statistical genetics. My work also includes viral genome-based epidemiological research and genomic breeding studies in livestock.

keywords：evolutionary biology, statistical genetics, virology

1. SGV-caller: SARS-CoV-2 genome variation caller. Heliyon 11（4）:e42613（2025）
2. The SARS-CoV-2 Lambda variant exhibits enhanced infectivity and immune resistance. Cell reports 38（2）110218（2022）
3. Genomic adaptation to polyphagy and insecticides in a major East Asian noctuid pest. Nature ecology & evolution 1（11）, 1747-1756 （2017）

伴侶動物、家畜、野生動物、動物園動物などを対象として、人間と動物の共生の観点から動物の福祉（アニマルウェルフェア）の向上を目指したフィールド研究を行っている。

My research aims for the improvement of animal welfare, especially the welfare of free-roaming cats, from the view point of the symbiotic relationships between humans and animals.

keywords：animal Welfare, human-animal relationships, free-roaming cats

1. Health status of 'community cats' living in the tourist area of the old town in Onomichi City, Japan. J Appl Anim Welf Sci 25: 338-354.（2022）
2. The effect of communal litter box provision on the defecation behavior of free-roaming cats in old-town Onomichi, Japan. Appl Anim Behav Sci 224: 104938.（2020）
3. Three-year route census study on welfare status of free-roaming cats in old-town Onomichi, Japan. J Appl Anim Welf Sci 21: 203-210.（2018）

甲殻類を中心とした無脊椎動物を対象に、フィールドワークと飼育実験から特に性差を示す形質（形態、行動、生理、遺伝子発現など）に着目して研究を行なっている。

My research group focuses on invertebrates, mainly decapod crustaceans, to understand sexual dimorphic characteristics such as morphology, behavior, physiology, and gene expression based on fieldwork and breeding experiments.

keywords：crustacean, sexual difference, aquaculture

1. Eyestalk transcriptome and methyl farnesoate titers provide insight into the physiological changes in the male snow crab, Chionoecetes opilio, after its terminal molt. Sci Rep 13:7204 (2023)
2. Dual roles of crustacean female sex hormone during juvenile stage in the kuruma prawn Marsupenaeus japonicus. Gen Comp Endocrinol 344:114374 (2023)
3. Impacts of methyl farnesoate and 20-hydroxyecdysone on larval mortality and metamorphosis in the kuruma prawn Marsupenaeus japonicus. Front Endocrinol 11:475 (2020)

自然物質およびそれらの農業および人間の健康保護への応用に焦点を当て、 具体的には糖尿病、肥満、 がんなどの慢性疾患の予防に関して研究しています。 さらに、 メタボロミクス研究を行い、さまざまな気象条件や環境圧力下での作物品種の反応メカニズムを解読しています。

Our group focuses on researching natural products and their applications in agriculture and human health protection, specifically in preventing chronic diseases such as diabetes, obesity, and cancer. In addition, we conduct metabolomic studies to decode the reaction mechanisms of crop varieties under various weather and environmental pressures.

keywords：natural products, rice, anti-chronic diseases, anticancer, metabolomics, environmental stress

1. Contribution of momilactones A and B to diabetes inhibitory potential of rice bran: Evidence from in vitro assays. Saudi Pharm J 27: 643-649 (2019)
2. Antioxidant, α-amylase and α-glucosidase inhibitory activities and potential constituents of Canarium tramdenum bark. Molecules 24: 605 (2019)
3. Anti-diabetes, anti-gout, and anti-leukemia properties of essential oils from natural spices Clausena indica, Zanthoxylum rhetsa, and Michelia tonkinensis. Molecules 27: 774 (2022)

ニワトリやウズラを用いて、エネルギー代謝の調節に関わる神経ペプチドの生理機能の解明を行っています。特に視床下部と脂肪組織、肝臓、生殖腺との関係性に着目しています。

My research focuses on the physiological functions of neuropeptides involved in the regulation of energy metabolism in chickens and quail. In particular, I focus on the relationship between the hypothalamus and adipose tissue, liver, and gonads.

keywords：neuroscience, metabolism, livestock, physiology, endocrinology

1. A novel hypothalamic factor, neurosecretory protein GM, causes fat deposition in chicks. Front  Physiol 12:747473 (2021)
2. Effect of stressors on the mRNA expressions of neurosecretory protein GL and neurosecretory protein GM in chicks. Front Physiol 13:860912 (2022)
3. Expression of mRNAs encoding hypothalamic small proteins, neurosecretory protein GL and neurosecretory protein GM, in the Japanese quail, coturnix japonica. Zoolog Sci 41:50-59 (2024)

イネの塩・アルカリストレス耐性機構を植物栄養学的観点から分子生理学的に研究しています。特に、同ストレス下におけるカリウム獲得機構に着目しています。

My research focuses on the molecular physiological mechanisms underlying saline-alkaline stress tolerance in rice, from the perspective of plant nutrition. In particular, I am interested in how rice acquires potassium from the rhizosphere under such stress conditions.

keywords：rice; saline-alkaline stress; potassium acquisition mechanisms; potassium transporter

1. Potassium transporter OsHAK17 may contribute to saline-alkaline tolerant mechanisms in rice（Oryza sativa）. J. Plant Res 137: 505-520 （2024）
2. Different rhizospheric pH conditions affect nutrient accumulations in rice under salinity stress, Plants 10: 1295（2021）
3. Characterization of Na+ exclusion mechanism in rice under saline-alkaline stress conditions. Plant Sci, 287: 110171（2019）

I have found that bacterial components or lipopolysaccharide or LPS can be translocated between organs and cause inflammation in the mammary gland. Therefore, my study investigates the relationship between mastitis and other diseases associated with LPS translocation in dairy animals.

keywords：mastitis, immunity, lipopolysaccharide, LPS, inflammation, ruminant, mammary gland

1. Mammary leukocytes function of endotoxin tolerant goat induced by intrauterine infusion of lipopolysaccharide. Innate Immun 31: 1-12 （2025）
2. Effect of intramammary lipopolysaccharide challenge after repeated intrauterine infusion of lipopolysaccharide on the inflammation status of goat mammary glands. Theriogenology 212: 104-110（2023）
3. Effect of repeated intrauterine infusion of lipopolysaccharides on mastitis in goats. Theriogenology 193: 87-92（2022）

I am driven to understand how plants function at a mechanistic level. Currently, I study how chemical priming induces plant stress memory and explore how three-dimensional vein networks form during development and in response to stress.

keywords：abiotic stress tolerance, Arabidopsis thaliana, auxin, chemical priming, plasmodesmata, Oryza sativa, plant stress memory, vascular development

1. Leaf vein patterning is regulated by the aperture of plasmodesmata intercellular channels. PLoS Biol 20, e3001781 (2022)
2. Of cells, strands, and networks: Auxin and the patterned formation of the vascular system. Cold Spring Harb Perspect Biol 13, (2021)
3. GAL4/GFP enhancer-trap lines for identification and manipulation of cells and tissues in developing Arabidopsis leaves. Developmental Dynamics 249, 1127–1146 (2020)